4v8s

Archaeal RNAP-DNA binary complex at 4.32Ang

Method: X-RAY DIFFRACTION

▼

1. Protein Identity and Related Structures Protein Identity & Related Structures

DNA-DIRECTED RNA POLYMERASE SUBUNIT K

OrganismNot specified

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB61_SULSH
Isoform	—
PDB entities	3
Chains and sequence ranges	Author chain AI； PDBConstruct 1–95； UniProt 1–95 Author chain BK； PDBConstruct 1–95； UniProt 1–95

RNA POLYMERASE SUBUNIT 13

OrganismNot specified

UniProt B8YB65

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB65_SULSH
Isoform	—
PDB entities	4
Chains and sequence ranges	Author chain AJ； PDBConstruct 1–104； UniProt 1–104 Author chain BQ； PDBConstruct 1–104； UniProt 1–104

DNA-DIRECTED RNA POLYMERASE SUBUNIT L

OrganismNot specified

UniProt B8YB62

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB62_SULSH
Isoform	—
PDB entities	5
Chains and sequence ranges	Author chain AM； PDBConstruct 1–92； UniProt 1–92 Author chain BL； PDBConstruct 1–92； UniProt 1–92

DNA-DIRECTED RNA POLYMERASE SUBUNIT N

OrganismNot specified

UniProt B8YB63

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB63_SULSH
Isoform	—
PDB entities	6
Chains and sequence ranges	Author chain AO； PDBConstruct 1–66； UniProt 1–66 Author chain BN； PDBConstruct 1–66； UniProt 1–66

DNA-DIRECTED RNA POLYMERASE

OrganismNot specified

UniProt B8YB55

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB55_SULSH
Isoform	—
PDB entities	7
Chains and sequence ranges	Author chain AR； PDBConstruct 1–1131； UniProt 1–1131 Author chain BB； PDBConstruct 1–1131； UniProt 1–1131

DNA-DIRECTED RNA POLYMERASE SUBUNIT D

OrganismNot specified

UniProt B8YB56

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB56_SULSH
Isoform	—
PDB entities	8
Chains and sequence ranges	Author chain AS； PDBConstruct 1–265； UniProt 1–265 Author chain BD； PDBConstruct 1–265； UniProt 1–265

RNA POLYMERASE SUBUNIT 7

OrganismNot specified

UniProt B8YB57

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB57_SULSH
Isoform	—
PDB entities	9
Chains and sequence ranges	Author chain AT； PDBConstruct 1–180； UniProt 1–180 Author chain BE； PDBConstruct 1–180； UniProt 1–180

RNA POLYMERASE SUBUNIT 4

OrganismNot specified

UniProt B8YB58

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB58_SULSH
Isoform	—
PDB entities	10
Chains and sequence ranges	Author chain AU； PDBConstruct 1–113； UniProt 1–113 Author chain BF； PDBConstruct 1–113； UniProt 1–113

RNA POLYMERASE SUBUNIT 8

OrganismNot specified

UniProt B8YB59

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB59_SULSH
Isoform	—
PDB entities	11
Chains and sequence ranges	Author chain AV； PDBConstruct 1–132； UniProt 1–132 Author chain BG； PDBConstruct 1–132； UniProt 1–132

DNA-DIRECTED RNA POLYMERASE

OrganismNot specified

UniProt B8YB53

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB53_SULSH
Isoform	—
PDB entities	12
Chains and sequence ranges	Author chain AW； PDBConstruct 1–880； UniProt 1–880 Author chain BA； PDBConstruct 1–880； UniProt 1–880

DNA-DIRECTED RNA POLYMERASE SUBUNIT P

OrganismNot specified

UniProt B8YB64

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB64_SULSH
Isoform	—
PDB entities	13
Chains and sequence ranges	Author chain AX； PDBConstruct 1–48； UniProt 1–48 Author chain BP； PDBConstruct 1–48； UniProt 1–48

;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ;

OrganismNot specified

UniProt B8YB54

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） DNA-DIRECTED RNA POLYMERASE SUBUNIT H × 1 （B8YB60） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB54_SULSH
Isoform	—
PDB entities	14
Chains and sequence ranges	Author chain AY； PDBConstruct 1–395； UniProt 1–395 Author chain BC； PDBConstruct 1–395； UniProt 1–395

DNA-DIRECTED RNA POLYMERASE SUBUNIT H

OrganismNot specified

UniProt B8YB60

State in the Current Structure

Assembly	Physical composition	Protein state	Molecular copy count	Associated components	Data consistency
1	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers
2	Protein–DNA	Heteromer	Protein 13 DNA 2	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3' × 1 ;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ; × 1 DNA-DIRECTED RNA POLYMERASE SUBUNIT K × 1 （B8YB61） RNA POLYMERASE SUBUNIT 13 × 1 （B8YB65） DNA-DIRECTED RNA POLYMERASE SUBUNIT L × 1 （B8YB62） DNA-DIRECTED RNA POLYMERASE SUBUNIT N × 1 （B8YB63） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB55） DNA-DIRECTED RNA POLYMERASE SUBUNIT D × 1 （B8YB56） RNA POLYMERASE SUBUNIT 7 × 1 （B8YB57） RNA POLYMERASE SUBUNIT 4 × 1 （B8YB58） RNA POLYMERASE SUBUNIT 8 × 1 （B8YB59） DNA-DIRECTED RNA POLYMERASE × 1 （B8YB53） DNA-DIRECTED RNA POLYMERASE SUBUNIT P × 1 （B8YB64） ;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ; × 1 （B8YB54） ZINC ION × 6 IRON/SULFUR CLUSTER × 1 MAGNESIUM ION × 1	Consistent with all polymers

Other States of the Same Protein in the Database

View Construct and Data Evidence

UniProt name	B8YB60_SULSH
Isoform	—
PDB entities	15
Chains and sequence ranges	Author chain AZ； PDBConstruct 1–84； UniProt 1–84 Author chain BH； PDBConstruct 1–84； UniProt 1–84

The page prioritizes protein identity, the current assembly, associated components, oligomeric state and cross-PDB links. Chain mapping and sequence ranges are retained as data evidence. Internal IDs, import timestamps and assembly operation expressions are maintenance fields and are not shown here.

▼

2. Structure Basics 2. Structure Basics

Entry ID entry_id	4v8s
Deposition date deposition_date	2012-07-12
Structure title title	Archaeal RNAP-DNA binary complex at 4.32Ang
Keywords keywords	TRANSFERASE-DNA COMPLEX, TRANSCRIPTION; TRANSFERASE/DNA
Experimental Method method	X-RAY DIFFRACTION

▼

3. Official assembly/model SAXS Official SAXS Profiles

This page reads only the latest assembly calculations. Every curve maps to an explicit PDB entry, official biological assembly and coordinate model.

4v8s__assembly_1__model_1

Assembly 1 · Model 1 · CRYSOL 4.1.3-1-20251215 (887e7ef)

Download this curve (.dat)

4v8s__assembly_1__model_1 | I(q)

100 plotted points; both axes use logarithmic scales.

4v8s__assembly_1__model_1 | P(r) · Pending

P(r) will be calculated and displayed separately for the same assembly/model.

Rg（Guinier）	49.74 Å
Rg (electron density)	49.49 Å
Total Rg	49.78 Å
Atom count	55761
Residues	3439
Excluded volume	495380 Å³
Maximum q	0.500 Å⁻¹

Assembly	Model	Structure unit	Oligomeric description	Status	CRYSOL	Actions
1	1	`4v8s__assembly_1__model_1`	pentadecameric （15）	Success	4.1.3-1-20251215 (887e7ef)	View Download
2	1	`4v8s__assembly_2__model_1`	pentadecameric （15）	Success	4.1.3-1-20251215 (887e7ef)	View Download

▶

4. Crystallography and Experiment 4. Crystallography & Experiment

▶

5. Entities and Polymers Entities & Polymers (18)

Entity 1 Entity 1 polymer

Description description	5'-D(TPCPTPTPAPTPAPCPTPCPTPAPTPCP)-3'
Molecular weight formula_weight	4180.7 kDa
Source method src_method	syn
Copy count entity_count	2

Polymer Polymer

Polymer type type	polydeoxyribonucleotide
Chain IDs strand_id	AC,BR
Atom count atom_count	111598

TCTTATACTCTATC

Entity 2 Entity 2 polymer

Description description	;5'-D(APTPAPGPAPGPTPAPTPAPAPGPAPTP APG)-3' ;
Molecular weight formula_weight	4994.3 kDa
Source method src_method	syn
Copy count entity_count	2

Polymer Polymer

Polymer type type	polydeoxyribonucleotide
Chain IDs strand_id	AD,BS

ATAGAGTATAAGATAG

Entity 3 Entity 3 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT K
Molecular weight formula_weight	10729.5 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AI,BK

MGLERDGILSQDLHFNEVFVSLWQNKLTRYEIARVISARALQLAMGAPALIDINNISSTDVISIAEEEFKRGVLPITIRRRLPNGKIILLSLRKS

Entity 4 Entity 4 polymer

Description description	RNA POLYMERASE SUBUNIT 13
Molecular weight formula_weight	12170.6 kDa
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AJ,BQ

MVSGMSTEEEKEGTNDEEVSEEREVEETSEEEFPKLSIQDIELLMKNTEIWDNLLNGKISVDEAKRLFEDNYKDYEKRDSRRKAKKAASKKVKKTKKKEKSVEG

Entity 5 Entity 5 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT L
Molecular weight formula_weight	10211.8 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AM,BL

MEIKILKSESNYLELEIEGEDHTLGNLIAGTLRKISGVSFASYYQPHPLTDKIIVKILTDGSIAPKDALLKAIETVRVMASHYIDEIKGLTK

Entity 6 Entity 6 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT N
Molecular weight formula_weight	7617.1 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AO,BN

MMIPIRCFTCGSLIADKWQPFITRVNAGENPGKVLDDLGVKRYCCRRMLLSHIDIISEVIHYTRPI

Entity 7 Entity 7 polymer

Description description	DNA-DIRECTED RNA POLYMERASE
Molecular weight formula_weight	127582.0 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AR,BB

MNELSSNLSIDERWKVIEAYFKSKGLVRQHLDSYNDFVRNKLQEIIDEQGEIPTEIPGLKVRLGKIRIGKPRVRESDRGEREISPMEARLRNLTYAAPLWLTMIPVENNIEAEPEEVYIGDLPIMLKSAIDPISQYTLDKLIEIGEDPKDPGGYFIVNGSERVIVTQEDLAPNRVLVDTGKTGSNITHTAKIISSTAGYRVPVTIERLKDGTFHVSFPAVPGKIPFVILMRALGILTDRDIVYAVSLDPEIQNELFPSLEQASSIANVDDALDFIGSRVAIGQKRENRIEKAQQIIDKYFLPHLGTSADDRRKKAYYLAYAISKVIELYLGRREPDDKDHYANKRLRLAGDLFASLFRVAFKAFVKDLTYQLEKSKVRGRKLALKALVRPDIVTERIRHALATGNWVGGRTGVSQLLDRTNWLSMLSHLRRVISSLARGQPNFEARDLHGTQWGRMCPFETPEGPNSGLVKNLALMAQIAVGINEKIVEKTLYEMGVVPVEEVIRRVTEGGEDQNEYLKWSKVILNGRLVGYYRDGEELAKKIRERRRKGEISDEVNVGHIVTDFINEVHVNCDSGRVRRPLIIVSNGNPLVTREDIEKLDSGSITFDDLVRQGKIEYLDAEEEENAYVALEPSDLTPEHTHLEIWSPAILGITASIIPYPEHNQSPRNTYQSAMAKQALGLYAANYQLRTDTRAHLLHYPQRPLVQTRALDIIGYTNRPAGNNAILAVISFTGYNMEDSIIMNRSSVERGMYRSTFFRLYSTEEVKYPGGQEDKIVMPEPGVRGYKGKEYYRLLEDNGVVSPEVEVKGGDVLIGKVSPPRFLQEFKELSPEQAKRDTSIVTRHGEMGIVDLVLITETAEGNKLVKVRVRDLRIPSIGDKFASRHGQKGVIGMLIPQVDMPYTVKGVVPDVILNPHALPSRMTLGQIMEGIAGKYAALSGNIVDATPFYKTPIEQLQNEILKYGYLPDATEVTYDGRTGQKIKSRIYFGVVYYQKLHHMVADKIHARARGPVQILTRQPTEGRAREGGLRFGEMERDCLIGFGTAMLLKDRLLDNSDRTTIYVCDQCGYIGWYDKNKNKYVCPIHGDKSNLFPVTVSYAFKLLIQELMSMIISPRLILEDRVGLSGGKGNE

Entity 8 Entity 8 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT D
Molecular weight formula_weight	30179.1 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AS,BD

MSINLLHKDDKRIDLVFEGYPLEFVNAIRRAAMLYVPVMSIDDVYFIENNSPLYDEILAHRLALIPFTSEEALDTYRWPEECIDCTENCEKCYTKIYIEAEALNEPKMLYSKDIKSEDPSIVPISGDIPIVLLGANQKISLEARLRLGYGKEHAKFIPVSLAIVRYYPKVEILGNCEKGATVCPEGVFELKDGKLSVKNELACTLCEECLRYCNGLIRISSVEDKYILELESVGSLKPERILLEAGKSIIRKIEELEKKLVEVIK

Entity 9 Entity 9 polymer

Description description	RNA POLYMERASE SUBUNIT 7
Molecular weight formula_weight	20328.7 kDa
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AT,BE

MYKLIKARSIVRIPPNEFGKPLNEIALNELRQQYQEKILKDLGLVLAILNVKTSEEGMLVFGDGATYHEVEFDMITYVPVVQEVVEGEVLQVDNYGVFVNLGPMDGLVHISQITDDTLKYDNVRGIIFGEKSKKVIQKGDKVRARVISVASTVTGRLPRIALTMRQPYLGKLEWITQAKK

Entity 10 Entity 10 polymer

Description description	RNA POLYMERASE SUBUNIT 4
Molecular weight formula_weight	12822.6 kDa
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AU,BF

MSSVYIVEEHYIPYSVAKKLLSDVIKSGSSSNLLQRTYDYLNSVEKCDAESAQKVVEELSSIISREDVRAVLASICPITPDEVRSILIMDSNRTYTSEDIQKIIDIIRKYIKS

Entity 11 Entity 11 polymer

Description description	RNA POLYMERASE SUBUNIT 8
Molecular weight formula_weight	15139.5 kDa
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AV,BG

MMESKAQEIILSCEINSIERGSLKNLSIIHMSCNDFNISFDIIDSINIFSQKEKVKAFISKNRLSYTNDDFCGHGYIVTELKDSSSNNGNRYITIISLFGLLVKIISNKESFLKIHQLNVMDHIYFCVKKNT

Entity 12 Entity 12 polymer

Description description	DNA-DIRECTED RNA POLYMERASE
Molecular weight formula_weight	99766.5 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AW,BA

MSEKNIKGIKFGILSPDEIRKMSVTAIITPDVYDEDGTPIEGSVMDPRLGVIEPGQKCPTCGNTLGNCPGHFGHIELVRPVIHVGFVKHVYEFLKATCRRCGRVKISEDEIEKYSRIYNAIKKRWPSAARRLTEYVKKTAMKAQVCPHCGEKQFKIKLEKPYNFYEERKEGVAKLTPSDIRERLEKVPESDVEILGYDPTTSRPEWMILTVLPVPPITIRPSIMIESGIRAEDDLTHKLVDIVRINERLKESIDAGAPQLIIEDLWDLLQYHVATYFDNEIPGLPPSKHRSGRPLRTLAQRLKGKEGRFRGNLSGKRVDFSSRTVISPDPNISIDEVGVPEIIARTLTVPERITPWNIEKLRQFVINGPDKWPGANYVIRPDGRRIDLRYVKDRKELASTLAPGYVVERHLTDGDVVLFNRQPSLHRISMMAHRVRVLKGLTFRLNLLVCPPYNADFDGDEMNLHVPQSEEAIAEAKEIMLVHKNIITPRYGGPIIGAAQDYISGAYLLTVKTTLLTKEEAQQILGVADVKIDLGEPAILAPREYYTGKQVVSAFLPKDFNFHGQANVSSGPRLCKNEDCPHDSYVVIKNGILLEGVFDKKAIGNQQPESILHWLIKEYSDEYGKWLMDNLFRVFIRFVELQGFTMRLEDVSLGDDVKKEIYNEIDRAKVEVDNLIQKYKNGELEPIPGRTLEESLENYILDTLDKLRSTAGDIASKYLDPFNFAYVMARTGARGSVLNITQMAAMLGQQSVRGERIKRGYMTRTLPHFKPYDISPEARGFIYSSFRTGLKPTELFFHAAGGREGLVDTAVRTSQSGYMQRRLINALSDLRAEYDGTVRSLYGEVIQVAYGDDGVFPMYSAHGKTVDVNRIFERVVGWKT

Entity 13 Entity 13 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT P
Molecular weight formula_weight	5607.0 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AX,BP

MAVYRCGKCWKTFTDEQLKVLPGVRCPYCGYKIIFMVRKPTIKIVKAI

Entity 14 Entity 14 polymer

Description description	;DNA-DIRECTED RNA POLYMERASE SUBUNIT A'' ;
Molecular weight formula_weight	43792.8 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AY,BC

MENVIDEKDKSYLEEKVKQASNILPQKIVEDLKNLISNKEVLVTRDEIDKIFDLAIKEYSEGLIAPGEAIGIVAAQSVGEPGTQMTLRTFHFAGIRELNVTLGLPRLIEIVDAKKVPSTPMMTIYLTDEYKHDKEKALEVARKLEYTKIENVVSSTSIDIASMSIILQLDNEMLKDKGVTVDDVKKAINRLKLGEFVIDESEGNTLNISFANIDSIAALFKLRDKILNTKIKGIKGIKRAIVQKKGDEYIILTDGSNLSGVLSVKGVDIAKVETNNIREIEEVFGIEAAREIIIREISKVLAEQGLDVDMRHILLVADVMTRTGVVRQIGRHGVTGEKNSVLARAAFEVTVKHLLDAAARGDVEEFKGVVENIIIGHPIKLGTGMVELTMRPILR

Entity 15 Entity 15 polymer

Description description	DNA-DIRECTED RNA POLYMERASE SUBUNIT H
Molecular weight formula_weight	9688.3 kDa
EC 编号 ec_number	2.7.7.6
Source method src_method	nat
Copy count entity_count	2

Polymer Polymer

Polymer type type	polypeptide(L)
Chain IDs strand_id	AZ,BH

MRGSSNRKIDPRIHYLVPKHEVLSIDEAYKILKELGIRPEQLPWIRASDPVARSINAKPGDIIRIIRKSQLYGEVVSYRYVISG

Entity 16 Entity 16 non-polymer

Description description	ZINC ION
Molecular weight formula_weight	65.4 kDa
Source method src_method	syn
Copy count entity_count	12

Entity 17 Entity 17 non-polymer

Description description	IRON/SULFUR CLUSTER
Molecular weight formula_weight	351.6 kDa
Source method src_method	syn
Copy count entity_count	2

Entity 18 Entity 18 non-polymer

Description description	MAGNESIUM ION
Molecular weight formula_weight	24.3 kDa
Source method src_method	syn
Copy count entity_count	2

▶

Unit-cell length a length_a	134.134 Å
Unit-cell length b length_b	199.416 Å
Unit-cell length c length_c	214.246 Å
Unit-cell angle α angle_alpha	90.00 °
Unit-cell angle β angle_beta	103.54 °
Unit-cell angle γ angle_gamma	90.00 °
Molecules per unit cell Z z_pdb	4

Space group space_group	P 1 21 1
Space-group number int_tables_number	4
Crystal system crystal_system	—

Resolution resolution	4.323 Å
Low-resolution limit resolution_low	50.424 Å
R factor r_factor	0.2926
R_work r_work	0.2917
R_free r_free	0.3105

Archaeal RNAP-DNA binary complex at 4.32Ang

1. Protein Identity and Related Structures Protein Identity & Related Structures

DNA-DIRECTED RNA POLYMERASE SUBUNIT K

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

RNA POLYMERASE SUBUNIT 13

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

DNA-DIRECTED RNA POLYMERASE SUBUNIT L

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

DNA-DIRECTED RNA POLYMERASE SUBUNIT N

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

DNA-DIRECTED RNA POLYMERASE

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

DNA-DIRECTED RNA POLYMERASE SUBUNIT D

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

RNA POLYMERASE SUBUNIT 7

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

RNA POLYMERASE SUBUNIT 4

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

RNA POLYMERASE SUBUNIT 8

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer

Other homooligomers

Heteromeric or nucleic-acid complex

DNA-DIRECTED RNA POLYMERASE

State in the Current Structure

Other States of the Same Protein in the Database

Monomer

Homodimer

Homotetramer